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In the pre-catalysis state, the S1 site would be occupied by the proximal SUMO2/3, a non-SUMO substrate protein, or by (acetyl-)CoA, as shown for ChlaDUB1 44

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From earlier work focused on muscular dystrophy conducted with former Penn Dental Medicine faculty member Elizabeth Barton, now at the University of Florida, the researchers believed that a particular form of IGF, a precursor of the protein that includes a separate component known as an e-peptide, was likely to stimulate regeneration better than mature IGF-1 that lacked the peptide

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