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After incubation, the WPs were centrifuged at 2,000 g for 10 min, and the supernatant was retained

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These three-stranded nucleic acid structures, composed of an RNA:DNA hybrid and a single-stranded DNA (ssDNA) loop, can form co-transcriptionally by invasion of the nascent transcript into the underwound DNA duplex behind the transcription complex, and act as a strong barrier to replication fork progression 10

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