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Funding This work was supported by NEI/NIH Intramural funding, Project EY000184 (R.R.C.), NIH EY026893 (C.S.d.P.), EY030447 (C.S.d.P.), EY025761 (A.J.St.L.), EY026891 (A.J.St.L.), EY032482 (A.J.St.L.)